I was going to come on to this post and argue that I didn't think Mark was into Juno "that way," but as I re-watched the basement scene with the two of them it is clear that he is feeling something. He is legitimately shocked by Juno's reaction to him leaving Vanessa, and expects that she would have been cool with it because she is into him. As she is ranting at him and he is sitting with his head almost in his hands commenting "I was so stupid" and "you're so young", its clear that he had thought she could have been something to lean on after leaving Vanessa, and she clearly is railing against that. I think he lets go of any romantic inclinations to her in that basement and in the follow up convo with Vanessa, but I can definitely see that something WAS there.
While Vanessa was pushing for a baby pretty hard and might have assumed it would fix all their marital issues, it's clear that Mark had been feeling that way for a while and was just waiting for a moment to ditch her. He was such an ass.
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