Perhaps it also aired in Europe and Australia and the like on their versions of Nickelodeon, hence they thought PAL would be a good idea?
What's also interesting is once they started having Hanho do the actual production layouts in season 2 (now working off key paintings that Jumbo did in house), Hanho had different teams of people working on different episodes, hence there was a lot of variation in both the art work and animation. Seasons 3 and 4 were pretty similar to each other overall, but season 2 was different from both 1 and 3/4. I suspect that part of it was the the result of Jumbo determining that their thin-lined (with occasional trademark wavy lines or squiggles) background art style was difficult for Hanho to copy after seeing that season 2 came out very hit or miss in that regard (whereas season 1 has some of the best ambiance ever seen in a cartoon). After all, Jinkins originally intended to do the actual characters with squiggly lines, as seen in Doug Can't Dance (inspired by R.O. Blechman), but Jinkins's animators pointed out that oversees in-betweeners would have difficulty replicating that so they went with the straight outlines instead. So for 3/4, they switched to pretty much straight (and thicker) outlines in the background to make it easier for Hanho.
One final thing I found interesting - the head of the background painting department for the Nickelodeon series, Michael Zodorozny, was not originally part of the Disney series. I suspect Disney wanted to try to "standardize" the show so that it looked more like other Disney cartoons, hence the new, garish color palette.
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